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Moreover, researchers fused TALEN to the Pseudomonas aeruginosa type III secretion system effector ExoS54 signal peptide and a nuclear localization signal, enabling direct cytosolic delivery and subsequent nuclear translocation of functional TALEN protein into human HeLa cells via bacterial injection, achieving targeted knockout of the Venus reporter gene [114]

355(a) and 331(d)

The confusion is understandable

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There is no good evidence for this, and it is worth being cautious about that framing