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This BtrG/GCT fold distinguishes these enzymes from others exhibiting cyclotransferase-like activity (Oakley et al., 2008)

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GCL is a heterodimeric enzyme composed of a glutamate-cysteine ligase catalytic subunit (GCLC), which confers enzymatic activity, and a glutamate-cysteine ligase modifier subunit (GCLM), which enhances GCLC efficiency and attenuates feedback inhibition by GSH