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Second, the cross-sectional nature of this study is a limitation which precludes our ability to make causal inferences about the observed associations

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First, the Hidden Markov Model (HMM) profile of the HMA domain PF00403 was downloaded from the Pfam database ( [32] and used as bait to conduct a BLAST search of the tobacco genome using the HMMER 3.0 program ( [33], and homologous proteins with an e-value 1e 4 in the output were screened as tobacco HIPP candidate proteins