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In 1962, through analysis of the radioactivity incorporated into collagen using a tritiated version of the amino acid proline, Stone and Meister discovered that vitamin C is used as a co-substrate by peptidyl-prolyl hydroxylase, an enzyme that catalyzes the selective modification of proline to hydroxyproline

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Furthermore, GHK-Cu's potential mechanism might involve the facilitation of copper uptake by cells, which might then be utilized by intracellular copper-dependent enzymes or directly activate transcription factors

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